Forage fish often make great migrations between their spawning, feeding and nursery grounds. Schools of a particular stock usually travel in a triangle between these grounds. For example, one stock of herrings have their spawning ground in southern Norway, their feeding ground in Iceland, and their nursery ground in northern Norway. Wide triangular journeys such as these may be important because forage fish, when feeding, cannot distinguish their own offspring. Fertile feeding grounds for forage fish are provided by ocean upwellings. Oceanic gyres are large-scale ocean currents caused by the Coriolis effect. Wind-driven surface currents interact with these gyres and the underwater topography, such as seamounts and the edge of continental shelves, to produce downwellings and upwellings. These can transport nutrients which plankton thrive on. The result can be rich feeding grounds attractive to the plankton feeding forage fish. In turn, the forage fish themselves become a feeding ground for larger predator fish. Most upwellings are coastal, and many of them support some of the most productive fisheries in the world. Regions of notable upwelling include coastal Peru, Chile, Arabian Sea, western South Africa, eastern New Zealand and the California coast. Capelin are a forage fish of the smelt family found in the Atlantic and Arctic oceans. In summer, they graze on dense swarms of plankton at the edge of the ice shelf. Larger capelin also eat krill and other crustaceans. The capelin move inshore in large schools to spawn and migrate in spring and summer to feed in plankton rich areas between Iceland, Greenland, and Jan Mayen. The migration is affected by ocean currents. Around Iceland maturing capelin make large northward feeding migrations in spring and summer. The return migration takes place in September to November. The spawning migration starts north of Iceland in December or January. The diagram on the right shows the main spawning grounds and larval drift routes. Capelin on the way to feeding grounds is coloured green, capelin on the way back is blue, and the breeding g ounds are red. In a paper published in 2009, researchers from Iceland recount their application of an interacting particle model to the capelin stock around Iceland, successfully predicting the spawning migration route for 2008. A gyre in oceanography is any large system of rotating ocean currents, particularly those involved with large wind movements. Gyres are caused by the Coriolis Effect; planetary vorticity along with horizontal and vertical friction, which determine the circulation patterns from the wind curl (torque). The term gyre can be used to refer to any type of vortex in the air or the sea, even one that is man-made, but it is most commonly used in oceanography to refer to the major ocean systems. Spawn is the eggs and sperm released or deposited, usually into water, by aquatic animals. As a verb, spawn refers to the process of releasing the eggs and sperm, also called spawning. Most aquatic animals, apart from aquatic mammals, reproduce through a process of spawning. Spawn consists of the reproductive cells (gametes) of aquatic animals, some of which will become fertilized and produce offspring. The process of spawning typically involves females releasing ova (unfertilized eggs) into the water, often in large quantities, while males simultaneously or sequentially release spermatozoa (milt) to fertilize the eggs. Most fish reproduce by spawning, and so do most other aquatic animals, including crustaceans such as crabs and shrimps, molluscs such as oysters and squid, echinoderms such as sea urchins and sea cucumbers, amphibians such as frogs and newts, other amphibious animals such as turtles, aquatic insects such as mayflies and mosquitoes, and corals (which are small aquatic animals and not plants). Fungi, such as mushrooms, are also said to "spawn" a white fibrous matter that forms the matrix from which they grow. There are many variations in the way spawning occurs, depending on sexual differences in anatomy, on how the sexes relate to each other, on where and how the spawn is released, and on whether or how the spawn is subsequently guarded.